Table 4.

Effect of sowing time on tannin (μg g⁻¹ DW), phytic acid (mg g⁻¹ DW), and trypsin inhibitor (units g⁻¹ DW) contents in mature grains of drought-susceptible (PBW 343) and drought tolerant (C 306) wheat genotypes grown under irrigated and rain-fed conditions. Values are mean ± standard deviation of three replicates; CD, critical difference; NS, not significant; One inhibitor unit is defined as the quantity of inhibitor which inhibits 50% of trypsin activity.

Year	Sowing time	Genotype	Tannin (μg g−1 DW)		Phytic acid (mg g−1 DW)		Trypsin inhibitors (units g−1 DW)
			Treatment		Treatment		Treatment
			Irrigated	Rain fed	Irrigated	Rain fed	Irrigated	Rain fed
Year I (2008)	Early sown	PBW 343	1608 ± 63.68	1382 ± 50.69	8.49 ± 0.50	6.00 ± 0.42	31.94 ± 0.98	33.80 ± 2.27
		C 306	1083 ± 48.03	1010 ± 18.03	7.85 ± 0.70	8.84 ± 0.68	45.83 ± 1.96	42.13 ± 1.13
	Timely sown	PBW 343	1840 ± 24.74	1689 ± 29.68	7.95 ± 0.62	9.38 ± 0.96	31.48 ± 3.97	27.31 ± 2.84
		C 306	1434 ± 16.00	1353 ± 33.79	8.00 ± 0.53	9.48 ± 1.02	40.28 ± 5.89	27.59 ± 6.57
	Late sown	PBW 343	1864 ± 20.87	1842 ± 19.00	9.58 ± 0.82	7.51 ± 0.45	29.63 ± 2.84	21.76 ± 2.27
		C 306	1767 ± 10.87	1470 ± 30.69	8.84 ± 0.40	6.62 ± 0.84	36.11 ± 5.97	32.87 ± 1.13
Year II (2009)	Early sown	PBW 343	1901 ± 24.17	1783 ± 32.50	7.70 ± 1.12	9.33 ± 1.28	34.26 ± 1.40	31.48 ± 1.57
		C 306	1426 ± 25.50	913 ± 12.50	7.56 ± 0.97	7.48 ± 0.64	46.30 ± 1.60	45.37 ± 1.52
	Timely sown	PBW 343	2020 ± 19.14	1426 ± 14.00	8.52 ± 0.36	9.26 ± 0.23	39.81 ± 1.13	31.02 ± 1.23
		C 306	1545 ± 29.48	1307 ± 19.67	9.74 ± 1.11	7.33 ± 0.59	41.20 ± 2.84	30.56 ± 0.98
	Late sown	PBW 343	1901 ± 22.03	1664 ± 27.57	10.00 ± 0.44	13.93 ± 0.28	30.56 ± 3.54	24.54 ± 1.50
		C 306	1248 ± 26.50	832 ± 20.88	9.22 ± 0.56	10.59 ± 0.90	39.81 ± 3.97	32.41 ± 1.45
			Tannin		Phytic acid		Trypsin inhibitors
			Year I	Year II	Year I	Year II	Year I	Year II
CD P ≤ 0.05	Sowing time		25.56	17.27	0.58	0.66	3.77	2.5
	Genotype		20.87	14.10	NS	0.54	3.08	2.05
	Treatment		20.87	14.10	0.48	0.54	3.08	2.05