Parameters
|
C
|
−0.02 |
m2
sµmol−
1
|
Slope of the linear relationship between I
fac and PPFD in the range 0–25 µmol m−
2 s−
1
|
|
35 |
Pa |
Reference atmospheric CO2 partial pressure |
d
|
1 |
µmol CO2 m−
2 leaf s−
1
|
y-intercept of the linear relationship between I
fac and PPFD in the range from 0–25 µmol m−
2 s−
1
|
g
b
|
300 |
mmol m−
2 s−
1
|
Leaf boundary layer conductance to water vapour |
g
fac
|
13.7 |
dimensionless |
Stomatal sensitivity coefficient |
g
min
|
76.2 |
mmol m−
2 s−
1
|
Minimum stomatal conductance to water vapour |
I
fac
|
0.5 |
dimensionless |
Coefficient representing the extent to which R
dark is inhibited in the light |
|
|
dimensionless |
Ratio between J
max and of plant grown at the reference temperature and at the reference CO2 partial pressure |
|
|
µmol CO2 g−
1 N s−
1
|
Slope of linear relationship relating N
pa to at the reference temperature and at the reference CO2 partial pressure |
K
c
|
19.42 |
Pa |
Michaelis-Menten constant for carboxylase activity of Rubisco |
K
o
|
14 300 |
Pa |
Michaelis-Menten constant for oxgenase activity of Rubisco |
O
i
|
21 000 |
Pa |
Internal leaf oxygen concentration |
p
1
|
−0.012 |
dimensionless |
Coefficient representing the extent to which J
fac is modified by the CO2 partial pressure during plant growth |
p
2
|
0.036 |
dimensionless |
Coefficient representing the extent to which J
fac is modified by the temperature during plant growth |
p
3
|
0.3192 |
µmol CO2 g−
1 N s−
1
|
Coefficient representing the effect of CO2 partial pressure during plant growth on k
3
|
p
4
|
0.94 |
dimensionless |
Coefficient representing the effect of growth temperature on entropy term for J
max and
|
R
|
8.314 |
J K−
1 mol−
1
|
Perfect gas constant |
R
fac
|
0.011 |
dimensionless |
Ratio between R
dark and at reference temperature |
SLA
|
|
m2 leaf g−
1 DM |
Specific leaf area |
α |
0.05 |
mol CO2 mol−
1 photon |
Apparent quantum yield of net photosynthesis at saturating CO2
|
ΔHaJmax
|
83 608 |
J mol−
1
|
Activation energy of J
max
|
ΔHaKc
|
65 800 |
J mol−
1
|
Activation energy of K
c
|
ΔHaKo
|
36 000 |
J mol−
1
|
Activation energy of K
o
|
ΔHaRdark
|
50 861 |
J mol−
1
|
Activation energy of R
dark
|
ΔHaVcmax
|
86 529 |
J mol−
1
|
Activation energy of
|
ΔHaτ
|
−28 990 |
J mol−
1
|
Activation energy of τ |
ΔHd |
200 000 |
J mol−
1
|
Deactivation energy |
|
660.42 |
J K−
1 mol−
1
|
Entropy term of J
max for plant grown at reference temperature |
|
654.24 |
J K−
1 mol−
1
|
Entropy term of for plant grown at reference temperature |
τ |
2 838 |
dimensionless |
Rubisco specificity factor at reference temperature |
Input Variables
|
C
a
|
|
Pa |
CO2 partial pressure in the ambient air |
C
g
|
|
Pa |
Atmospheric CO2 partial pressure during preceding month of plant growth |
h
s
|
|
dimensionless |
Leaf surface relative humidity |
PPFD
|
|
µmol m−
2 s−
1
|
Photosynthetic photon flux density |
T
k
|
|
K |
Air temperature. In our analysis T
k = T
g
|
T
g
|
|
K |
Mean air temperature during preceding month of plant growth |
T
r
|
293.16 |
K |
Reference temperature for metabolic activity |
|
|
|
|
Output variables
|
A
n
|
|
µmol m−
2 s−
1
|
Net photosynthesis |
C
i
|
|
Pa |
Internal CO2 partial pressure |
C
s
|
|
Pa |
Leaf surface CO2 partial pressure |
g
s
|
|
mmol m−
2 s−
1
|
Stomatal conductance to water vapor |
k
2
|
|
Pa |
Intermediate variable synthesizing the Rubisco affinity for CO2
|
J
|
|
µmol m−
2 s−
1
|
Light dependence of the rate of electron transport |
|
|
dimensionless |
J
fac acclimated to CO2 during plant growth |
|
|
dimensionless |
J
fac acclimated to temperature during plant growth |
|
|
dimensionless |
J
fac acclimated to CO2 and to temperature during plant growth |
J
max
|
|
µmol m−
2 s−
1
|
Potential rate of RuBP regeneration |
|
|
µmol m−
2 s−
1
|
Potential rate of RuBP regeneration at reference temperature |
k3
|
|
µmol CO2 g−
1 N s−
1
|
Slope of linear relationship relating N
pa to
|
|
|
µmol CO2 g−
1 N s−
1
|
Slope of linear relationship relating N
pa to acclimated to CO2 during plant growth |
N
a
|
|
g N m−
2 leaf |
Leaf N content per leaf area |
N
ac
|
|
g N m−
2 leaf |
Leaf N content per leaf area when W
c equals W
j
|
N
pa
|
|
g N m−
2 leaf |
Leaf photosynthetic N content per leaf area |
Np
ac
|
|
g N m−
2 leaf |
Leaf photosynthetic N content per leaf area when W
c equals W
j
|
R
dark
|
|
µmol m−
2 s−
1
|
Leaf dark respiration rate |
|
|
µmol m−
2 s−
1
|
Leaf dark respiration rate at reference temperature |
R
day
|
|
µmol m−
2 s−
1
|
Leaf respiration rate from processes other than photorespiration |
|
|
µmol m−
2 s−
1
|
Maximum carboxylation rate of Rubisco |
|
|
µmol m−
2 s−
1
|
Maximum carboxylation rate of Rubisco at reference temperature in the absence of any deactivation as a result of high temperature |
W
c
|
|
µmol m−
2 s−
1
|
Rubisco-limited photosynthetic rate |
W
j
|
|
µmol m−
2 s−
1
|
RuBP regeneration limited photosynthetic rate through electron transport |
|
|
dimensionless |
Temperature dependence of J
max or
|
|
|
dimensionless |
Temperature dependence of
|
|
|
dimensionless |
Temperature dependence of J
max
|
|
|
dimensionless |
Temperature dependence of K
c, K
o, τ, or R
dark
|
|
|
dimensionless |
Temperature dependence of K
c
|
|
|
dimensionless |
Temperature dependence of K
o
|
|
|
dimensionless |
Temperature dependence of τ |
|
|
dimensionless |
Temperature dependence of R
dark
|
Γ* |
|
dimensionless |
CO2 compensation point in the absence of mitochondrial respiration |
|
|
J K−
1 mol−
1
|
Entropy term acclimated to temperature during plant growth |
|
|
J K−
1 mol−
1
|
Entropy term of J
max acclimated to temperature during plant growth |
|
|
J K−
1 mol−
1
|
Entropy term of acclimated to temperature during plant growth |