Table 1. Symbol definitions.

Symbol	Definition
	Haploid population size
	Genomic beneficial mutation rate
	Total genetic map length
	Selective advantage of beneficial mutations
	Probability of fixation of a beneficial mutation
	Genomic rate of fixation of beneficial mutations
	Heritable variance in log fitness in the population
	Values of and in the absence of interference
	Expected time for a pair of neutral lineages to coalesce

The definitions of the main symbols used in the text. , , , , , and are population parameters, and , , , and are variables. In addition, we use to denote the expectation of a variable taken over a distribution of selective coefficients , and to denote the expectation over possible genetic backgrounds.