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The predominant conformation of the His64 side chain is neutral and in the “inward” position. The 2Fo−Fc nuclear map (contoured at 1.5σ) is shown in blue with the positive omit Fo−Fc nuclear density map (contoured at 3.0σ) in yellow, indicating the D atom density for the flipped neutral histidine conformation. Waters are as labeled, H atoms on the His imidazole ring are omitted for clarity.
