Table 2. Comparison of full and constrained maximum likelihood binary-state speciation and extinction (BiSSE) models.
Model | constraints | lambda0 | lambda1 | mu0 | mu1 | q01 | q10 | parameters | lnLik | AIC |
full | None | 0.0919287 | 0.0916504 | 0.0196976 | 0.0000042 | 0.0015639 | 0.0011354 | 6 | −775.51 | 1563.0 |
constraint 1 | lambda0 = lambda1 | 0.0917114 | 0.0917114 | 0.0195089 | 0.0000005 | 0.0015538 | 0.0011399 | 5 | −775.51 | 1561.0 |
constraint 2 | mu0 = mu1 | 0.0782689 | 0.0915912 | 0.0000147 | 0.0000147 | 0.0015438 | 0.0011227 | 5 | −775.59 | 1561.2 |
constraint 3 | q01 = q10 | 0.0917787 | 0.0917094 | 0.0197393 | 0.0000002 | 0.0013342 | 0.0013342 | 5 | −775.61 | 1561.2 |
constraint 4 | lambda0 = lambda1, mu0 = mu1 | 0.0858881 | 0.0858881 | 0.0000149 | 0.0000149 | 0.0015973 | 0.0010989 | 4 | −777.70 | 1563.4 |
constraint 5 | lambda0 = lambda1, q01 = q10 | 0.0917360 | 0.0917360 | 0.0197087 | 0.0000034 | 0.0013375 | 0.0013375 | 4 | −775.61 | 1559.2 |
constraint 6 | mu0 = mu1, q01 = q10 | 0.0781298 | 0.0916917 | 0.0000117 | 0.0000117 | 0.0013121 | 0.0013121 | 4 | −775.75 | 1559.5 |
constraint 7 | lambda0 = lambda1, mu0 = mu1, q01 = q10 | 0.0857970 | 0.0857970 | 0.0000003 | 0.0000003 | 0.0013039 | 0.0013039 | 3 | −777.92 | 1561.8 |
constraint 8 | q01 = 0 | 0.0815919 | 0.1616154 | 0.0000076 | 0.0960942 | 0.0000000 | 0.0019102 | 5 | −778.31 | 1566.6 |
constraint 9 | q10 = 0 | 0.1898625 | 0.0943962 | 0.1408593 | 0.0000000 | 0.0036345 | 0.0000000 | 5 | −820.50 | 1651.0 |
Trait 0 lacks adhesive toepads; trait 1 possesses adhesive toepads. Lambda = trait specific speciation rates; mu = trait specific extinction rates; q = transition rate parameters. Constrained models are compared using the Akaike Information Criterion (AIC). The models with the lowest AIC scores are in bold.