Table 2. Comparison of full and constrained maximum likelihood binary-state speciation and extinction (BiSSE) models.

Model	constraints	lambda0	lambda1	mu0	mu1	q01	q10	parameters	lnLik	AIC
full	None	0.0919287	0.0916504	0.0196976	0.0000042	0.0015639	0.0011354	6	−775.51	1563.0
constraint 1	lambda0 = lambda1	0.0917114	0.0917114	0.0195089	0.0000005	0.0015538	0.0011399	5	−775.51	1561.0
constraint 2	mu0 = mu1	0.0782689	0.0915912	0.0000147	0.0000147	0.0015438	0.0011227	5	−775.59	1561.2
constraint 3	q01 = q10	0.0917787	0.0917094	0.0197393	0.0000002	0.0013342	0.0013342	5	−775.61	1561.2
constraint 4	lambda0 = lambda1, mu0 = mu1	0.0858881	0.0858881	0.0000149	0.0000149	0.0015973	0.0010989	4	−777.70	1563.4
constraint 5	lambda0 = lambda1, q01 = q10	0.0917360	0.0917360	0.0197087	0.0000034	0.0013375	0.0013375	4	−775.61	1559.2
constraint 6	mu0 = mu1, q01 = q10	0.0781298	0.0916917	0.0000117	0.0000117	0.0013121	0.0013121	4	−775.75	1559.5
constraint 7	lambda0 = lambda1, mu0 = mu1, q01 = q10	0.0857970	0.0857970	0.0000003	0.0000003	0.0013039	0.0013039	3	−777.92	1561.8
constraint 8	q01 = 0	0.0815919	0.1616154	0.0000076	0.0960942	0.0000000	0.0019102	5	−778.31	1566.6
constraint 9	q10 = 0	0.1898625	0.0943962	0.1408593	0.0000000	0.0036345	0.0000000	5	−820.50	1651.0

Trait 0 lacks adhesive toepads; trait 1 possesses adhesive toepads. Lambda  =  trait specific speciation rates; mu  =  trait specific extinction rates; q  =  transition rate parameters. Constrained models are compared using the Akaike Information Criterion (AIC). The models with the lowest AIC scores are in bold.