Table 1 . Chromosome associations in PMCs of hybrids at diakinesis and metaphase I as revealed by FISH.

		Chromosome associations (ranges)
		I				II							III	IV
Hybrids	2n	Total	IA	IB	IC	Total	IIA-A	IIB-B	IIC-C	IIA-B	IIA-C	IIB-C	Total	Total	PMCs with multivalent (%)	Total PMCs
A.B	18	8.26 (3–16)	4.74a (2–9)	3.53b (1–7)	—	2.13 (0–5)	0.53b (0–3)	0.23c (0–2)	—	1.38c (0–5)	—	—	0.75 (0–3)	0.70 (0–2)	86.89	61
A.C	19	4.86 (1–13)	2.64b (0–8)	—	2.23a (0–5)	5.77 (3–9)	1.18a (0–3)	—	1.14b (0–3)	—	3.45c (0–7)	—	0.86 (0–3)	—	83.05	22
B.C	17	9.43 (3–13)	—	4.72a (1–7)	4.72d (1–9)	2.10 (0–5)	—	0.30b,c (0–2)	0.57c (0–2)	—	—	1.23c (0–5)	0.65 (0–3)	0.30 (0–1)	74.17	120
A.C.B	27	5.27 (2–8)	0.64c (0–2)	4.32a (2–7)	0.32c (0–2)	10.32 (2–12)	1.73d (0–3)	0.45b,a (0–1)	1.14b (0–3)	1.00c (0–3)	4.50b (1–8)	1.50c (0–5)	0.41 (0–1)	—	40.91	22
BB.A	26	7.21 (3–10)	5.65d(1–9)	1.56c (0–6)	—	8.12 (6–10)	0.65b (0–2)	5.35d (2–7)	—	2.12b (0–5)	—	—	0.85 (0–2)	—	67.65	34
CC.A	28	5.91 (3–10)	5.18a,d (2–9)	—	0.74c (0–4)	10.38 (8–12)	0.32c (0–2)	—	6.29a (2–9)	—	3.76c (0–8)	—	0.44 (0–1)	—	44.12	34
CC.B	26	6.40 (4–14)	—	4.80a (3–7)	1.60b (0–7)	8.70 (6–10)	—	0.70a (0–1)	6.70a (5–8)	—	—	1.20c (0–3)	0.80 (0–2)	—	70	10

I, univalent; II, bivalent; III, trivalent; IV, quadrivalent. I^A, I^B, and I^C indicate univalents belonging to the A, B, and C genomes, respectively; II^A-A, II^B-B, and II^C-C indicate autosyndetic bivalents formed between two chromosomes of A, B, or C genomes in A.B, A.C, B.C, and A.C.B hybrids, but II^B-B in BB.A and II^C-C in CC.A and CC.B indicate homologous bivalents; II^A-B, II^A-C, and II^B-C indicate allosyndetic bivalents formed between A and B chromosomes, A and C chromosomes, and B and C chromosomes, respectively.

^a,b,c,dGroups significantly different by χ² -test, p < 0.05.