Neighbour-joining tree obtained using a distance analysis with a general time reversible (GTR) substitution model on the partial 28S nrRNA gene alignment (812 nucleotides including alignment gaps) as implemented in PAUP v. 4.0b10 (Swofford 2003). Novel species are indicated in a red font and the orders are indicated on the right-hand side of the figure. The scale bar indicates the number of substitutions per site and the bootstrap support values (based on 1 000 replicates) are shown by colour-coded dots for values > 79 % (see legend on figure). The tree was rooted to species of the order Peronosporales.