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Mutations in AAA1 and AAA3 cause defects in nuclear segregation. To monitor nuclear segregation, the number of binucleate mitotic cells was determined after growth at 16°C for 16 h. The mean and standard error of the proportion are shown (n > 200). Walker A mutants (predicted to block nucleotide binding) are shaded with gray, whereas Walker B mutants (predicted to block nucleotide hydrolysis) are shaded with black.
