Fitness is maximized for negative maternal inheritance in stochastic environments: in the noisy reference environment (top) and in a stochastic environment with δ = 10 (bottom). The optimal value of the maternal effect coefficient m depends on the strength of environmental autocorrelation, ρτ. Darker grey indicates larger ρτ; values are (from light to dark) , , , , for which fitness is maximized at m =−0.2, −0.2, 0 and 0 (top) and m =−0.4, −0.4, − 0.3 and −0.3 (bottom), respectively. Parameters are as in figure 1. Circles represent the results of numerical simulations at intervals of 0.1 in m.