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. 2012 Jul 30;(212):63–79. doi: 10.3897/zookeys.212.3254
1 Head very wide and flat, posterior margin of the head not excavate, without posterior furrows. Epistomal lateral edge not or less oblique from the antennal basis to the dorsal articulation of mandible. Mouthparts occupant more than a half of the front margin of the head. Antennae not longer than the head. Lateral chitinous thorns of tergum IX gently curved backwards 2
Head wide and flat, with distinctly furrowed postgenae at the posterior margin, forming a separate swelling on the occipital part of the head. Mouthparts more protruding, lateral border of epistomal margin distinctly oblique from the antennal basis to the insertion of mandible. Mouthparts complex slender, antennae as long as the whole head. Lateral thorn of the tergum IX strongly bent backwards 3
2 Antennae slender, second joint distinctly longer than the first one, apical joint thin five times longer than wide at the basis. Lateral border of parietale a little swollen in correspondence of the stemmata. Epistomal front margin moderately oblique towards the mandibular basis (Figs 24–25). Urogomphi well curved and apically a little converging to the median plane (Figs 26–27). Basal tooth with minor spine directed outwards and far from the apex. Spiracular process small and little pronounced (Figs 36–38). Conical appendage very large at the basis, distinctly setose around the apical part, chitinous apex short (Fig. 37) Cucujus cinnaberinus (Scop.)
Antennae very short, second joint longer like the first one. Apical antennomere four times as wide as at the basis. Head robust and not swollen in the stemmata area (Figs 28–29). Front epistomal margin straight from antennal basis to mandibular insertion. Maximum head width behind the stemmata. Urogomphi well curved but apically not converging to the median plane (Figs 30–31). Basal tooth with less distant apical spines. Conical appendage slender and of different shape (Figs 39–41). Spiracular process inconspicuous Cucujus haematodes Er.
3 Head posteriorly sinuate and marked by very long antennae. Second antennomere a little longer than the first one. Apical antennomere six times longer than wide at the basis. Epistomal front margin strongly oblique towards the mandibular insertion. Frontal suture less sinuate than in previous species (Figs 32–33). Urogomphi strong and a little converging at the end, apex with three evident macrochaetae (Figs 34–35). Basal tooth of normal shape and slender. Spiracular process distinctly protrudent backwards (Figs 42–43). Lateral thorn of tergum IX strongly bent to the rear, forming an angle of ninety degrees (Fig. 44) Cucujus tulliae sp. n.