Fig. 2.

Basic structure of the hippocampal delay line model of adaptive timing as proposed by Zipser (1986). Each synapse introduces a delay; the total delay from activation of the first element in the delay-line to the last element is a direct function of the number of sequential synapses. Taps from the delay-line units send timing information to higher-order processing units. NMR(t) is the nictitating membrane response produced by US, which acts as teaching signal. unmodified, modified synapse. CS input initiates sequential propagation of signal through a delay-line