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Schematic of the DNA winding pattern along two neighbouring nucleosomes in the two-angle model. (a) View down the NCP axis ai. (b) View down the linker direction ti−1. The angle α is the dihedral angle (ti−1, ai, ti) and β the dihedral angle (ai−1, ti−1, ai) standing for the twist (modulo 2π) of the DNA linker.
