Table 1. Intercolony variation in seven training parameters for experiment 2.

Colony	D3	D4	D6	D8	D9	D10	Kruskal-Wallis (X 2) p-value
	mean ± S.E. (median)
a) Start time of training	11:57±00:25 (11:40)	12:52±00:35 (12:29)	14:07±00:07 (14:12)	13:46±00:39 (13:52)	13:01±00:34 (13:11)	12:44±00:28 (13:11)	(10.00) 0.075
Initial Learning
b) Number of bouts	4.13±0.35 (4)	4.87±0.39 (5)	3.8±0.09 (4)	3.73±0.42 (3)	3.67±0.23 (4)	3.73±0.53 (3)	(7.52) 0.185
c) bout duration/secs	246.7±17.5 (260)	267.8±26.5 (235)	287.9±9.8 (260)	354.7±61.3 (234)	237.6±17.6 (222)	271.1±34.7 (253)	(2.53) 0.771
Reversal Learning
d) Number of bouts	4.27±0.49 (4)	5.73±0.57 (6)	4.93±0.51 (5)	5.14±0.40 (5)	4.47±0.36 (4)	4.67±0.65 (5)	(5.52) 0.356
e) bout duration/secs	251.3±26.9 (246)	260.0±26.6 (241)	212.0±20.3 (190.5)	244.5±26.7 (207.5)	227.1±14.6 (195)	224.7±24.1 (199)	(4.90) 0.428
Number of Landings
f) yellow lands (initial)	29.9±2.0 (28)	37.4±2.8 (33)	25.3±2.2 (27)	31.4±2.2 (30)	36.1±2.5 (36)	30.9±2.7 (29)	(12.41) 0.029
g) blue lands (reversal)	32.1±3.1 (35)	45.7±4.0 (43)	38.0±3.4 (40.5)	42.1±2.7 (43.5)	44.0±3.2 (42)	38.7±4.0 (38)	(7.72) 0.173

(a) The average time when training started for bees in each of the six colonies. The average number (b) and duration (c) of bouts performed by bees in the initial training phase. The average number (d) and duration (e) of bouts performed by bees in the reversal training phase. The average number of landings made by a bee on rewarding flowers, yellow landings during initial training (f) and blue landings in reversal training (g), during the 100 flower choices (including the first time it probed a rewarding flower for the first time). In all cases the mean (± S.E.) and median values are given for each colony. The results from Kruskal-Wallis tests indicate whether variation among colonies is significant (statistically significant p-values are shown in bold).