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Phylogram showing the occurrence of modular vs. nonmodular social systems in (Asian) colobines. Pygathrix is classified as semimodular because the evidence for modularity in this genus is still equivocal (two species have been tentatively described as modular and one as mm–mf (see Table I). Phylograms were constructed in MacClade and phylogenies are based on Perelman et al. (2011) and Roos et al. (2011).
