Table 1. Parameters varied (variables) as a result of model-cycle testing and the parameterization resulting from the POM testing.

Parameter Ref	Function	Value 2003/2009	Value after POM
V1	Male minimum reproductive age (days)	40	30
V2	Female minimum reproductive age (days)	20	23
V3	A multiplier to get a quality score from area (e.g. 1.5×minimum home range)	2.0	2.1
V4	Minimum female territory radius (m)	8	8
V5	Maximum female territory radius (m)	16	8
V6	Minimum male territory radius (m)	12	9
V7	Maximum male territory radius (m)	20	23
V8	Age difference needed before a male can ‘evict’ a younger male (days)	0	30
V9	Additional probability of mortality on dispersal	0	0.055
V10	Daily unattributed mortality probability	0.003	0.0025
V11**	The date in autumn at which reproduction cannot be started (day)	273	230
V12	The probability of moving if there are no females over-lapping a male’s territory	NA	0.0505
V13	Threshold number of voles in a territory for density dependence effects	1	4
V14	The temperature at which grass is assumed to grow (triggers breeding if achieved for 7 consecutive days) (°C)	5	3.552
V15	The date before which breeding is impossible regardless of temperature (day)	70	80
V16*	The number of consecutive days a vole can disperse without dying (days)	5	infinite
V17*	Probability of infanticide attempt	100%	100%

^*V1–V15 were subsequently utilized in the sensitivity analysis, V16–V17 were found to be insensitive and therefore the effect of varying these was not reported.

^**V11 is a climate dependent parameter, fitted to Finnish conditions from Pattern Set 1. It should be adjusted when applied to other regions.