Anatomy of Pollia condensata fruit. (A) SEM image of the fruit surface showing smooth cuticular layer. (B) TEM cross-section showing three distinct tissue zones: (1) an outer epicarp of 3–4 layers of thick-walled cells, (2) an intermediate region of 2–3 layers of tanniniferous cells, and (3) a zone of thin-walled cells. (C) TEM of a single thick-walled cell from layer 1. (D) TEM of the cellulose microfibrils that constitute the thick cell wall in layer 1. The red lines highlight the twisting direction of the microfibrils. (E) (Left) Scheme showing a wedge of an LH helicoid with the arched pattern exposed on the oblique face (adapted from ref. (24)). (Right) Circularly polarized beams of light, with the wave vector of the light. The handedness of the transmitted and reflected light depends on the handedness of the helicoid. Here, light transmitted through the structure ( pointing down) is LH circularly polarized, while the reflected light ( pointing up) is RH. (F) 3D representation of the orientation of cellulose microfibril assembly and a transmitted circularly polarized beam.