Phylogenetic relationships between species and populations estimated by jointly analysing COI and rpS9 loci in *beast. Branches are coloured by species as shown in the key and the outgroup is Anopheles koliensis (black). Support values are posterior probabilities, and the scale bar shows time in millions of years (based on the molecular clock used—2.3% per million years for mtDNA; nuclear DNA estimated by the program). The error bar on the node at which An. farauti and An. hinesorum split is the 95% HPD (height) and shows the error associated with the estimate of the timing of divergence between the species. The shaded area shows the 95% HPD (height) of the putative introgression event based on the mtDNA beast phylogeny (node e). Abbreviations include: SI, Solomon Islands; MI, Manus Island; PP, Papuan Peninsula; n.NG, northern New Guinea; s.NG, southern New Guinea; s.SI, southern Solomon Islands; and n.SI, northern Solomon Islands.