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Biosynthetic pathway of HS/heparin. Enzymes shown in gray are not expressed in CHO-S host cells. NDST, N-deacetylase/N-sulfotransferase; GLCE, glucuronyl C5-epimerase; HS2ST, heparan sulfate 2-O-sulfotransferase; HS6ST, heparan sulfate 6-O-sulfotransferase; HS3ST, heparan sulfate 3-O-sulfotransferase.
