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Proposed three endogenous GA 20-oxidations in Arabidopsis. Different derivatives of C20-GAs can be oxidized by endogenous 20-oxidases to produce C19-GAs. The early 13-hydroxylation pathway leads to GA20; the non-3,13-hydroxylation pathway leads to GA9; and the early 3-hydroxylation pathway leads to GA4. GA9 can be 13-hydroxylated and 3β-hydroxylated to give GA20 and GA4. GA4 can be 13-hydroxylated and GA20 can be 3β-hydroxylated to give to GA1.
