Table 3. Variation partitioning of environmental and spatial effects on web-spider metacommunity composition.

Fractions of variation	R2	R2 adj	F	P
[a+b] Environmental + shared	0.30	0.12	1.68	0.005
[b+c] Spatial + shared	0.19	0.07	1.56	0.01
[a+b+c]	0.43	0.15	1.52	0.005
[a] Only environmental		0.08	1.40	0.025
[b] Environment spatially structured		0.05
[c] Only spatial		0.02	1.19	0.23
[d] Residual		0.85

Environmental variables: number of bushes, land use surrounding the patches, shape of patches. Spatial variables: two PCNM variables.