A, isometric force development and force response to four length steps of different sizes at 200 ms intervals. Each step is followed, after 50 ms, by a step in the opposite direction. From top to bottom traces are fibre length change, hs length change and force. Fibre length 5.01 mm; CSA 0.032 mm2; 5°C. B, sections of the records at the time of the steps, superimposed and on an expanded time scale. C, T1 relations for the fibre (•) and for the half-sarcomere (○) obtained from the records in B.