Table 4. Means, ± S.D., range, within-individuals variability (C.V._w) and between-individuals variability (C.V._b) for the four acoustic features analyzed from aggressive sounds produced by 14 Amphiprion akallopisos.

Acoustic variables	Overall Mean ± S.D. (range)	C.V.w (Mean)	C.V.w (range)	C.V.b	C.V.b/C.V.w	H*	p-value
Dominant frequency (Hz)	663±199 (346–1207)	0.10	0.07–0.21	0.30	2.73	1577	<0.001
Pulse duration (ms)	13.4±3.9 (3.8–22.9)	0.10	0.06–0.18	0.29	2.90	1614	<0.001
Pulse period (ms)	75.9±13.4 (32.4–121.9)	0.15	0.09–0.21	0.23	1.53	203.9	<0.001
Number of pulses per sound	3.9±2.2 (2–14)	0.50	0.30–0.68	0.56	1.12	25.55	0.0195

^* Results of Kruskall-Wallis test (df = 13, n = 1818) comparing differences between 14 individuals of A. akallopisos for each acoustic feature. Note that these values were calculated based on acoustic data obtained from Colleye et al. (2009). n, number of pulses analyzed.