Table 5. Means, ± S.D., range, within-individuals variability (C.V._w) and between-individuals variability (C.V._b) for the seven acoustic features analyzed from submissive sounds produced by 9 Amphiprion frenatus.

Acoustic variables	Group number	Overall Mean ± S.D. (range)	C.V.w (Mean)	C.V.w (range)	C.V.b	C.V.b/C.V.w	H*	p-value
Dominant frequency (Hz)	1	631±135 (431–1036)	0.10	0.09–0.15	0.21	1.90	213.7	<0.001
	2	593±139 (431–862)	0.10	0.08–0.12	0.22	2.20	251.0	<0.001
	3	552±105 (345–776)	0.09	0.08–0.11	0.19	2.11	239.9	<0.001
Pulse duration (ms)	1	7.4±2.1 (3.8–11.2)	0.10	0.09–0.11	0.28	2.80	232.4	<0.001
	2	8.5±2.8 (3.6–15.1)	0.10	0.09–0.11	0.34	3.40	250.8	<0.001
	3	9.1±2.8 (4.5–13.9)	0.08	0.06–0.12	0.31	3.87	233.2	<0.001
Pulse period (ms)	1	10.7±2.0 (6.7–15.4)	0.10	0.08–0.12	0.19	1.90	143.8	<0.001
	2	11.9±2.5 (7.4–17.6)	0.10	0.08–0.11	0.21	2.10	167.4	<0.001
	3	12.7±2.6 (8.2–18.4)	0.08	0.05–0.09	0.21	2.62	176.2	<0.001
Number of pulses per sound	1	3.1±0.6 (2–5)	0.20	0.19–0.23	0.21	1.05	5.36	ns
	2	3.4±0.9 (2–6)	0.26	0.21–0.31	0.27	1.04	2.59	ns
	3	3.1±0.8 (2–5)	0.25	0.22–0.29	0.27	1.08	12.86	<0.01
Sound duration (ms)	1	29.2±8.4 (9.8–48.2)	0.22	0.18–0.24	0.29	1.32	59.24	<0.001
	2	36.9±14.1 (16.6–72.4)	0.28	0.22–0.33	0.38	1.36	25.94	<0.001
	3	36.6±14.1 (15.0–73.8)	0.24	0.13–0.35	0.38	1.58	49.90	<0.001
Sound period (ms)	1	176.4±36.9 (92.2–295.3)	0.18	0.16–0.21	0.21	1.17	16.53	<0.01
	2	206.3±40.9 (135.6–280.9)	0.15	0.12–0.18	0.20	1.33	21.07	<0.001
	3	201.7±31.5 (118.3–266.3)	0.13	0.08–0.16	0.16	1.23	16.17	<0.01
Number of sounds per train	1	3.3±1.5 (2–9)	0.39	0.28–0.53	0.44	1.13	4.36	ns
	2	3.2±1.3 (2–7)	0.35	0.30–0.41	0.41	1.17	4.93	ns
	3	2.6±0.8 (2–5)	0.23	0.18–0.27	0.30	1.30	14.39	<0.01

^* Results of Kruskal-Wallis test (df = 2, n = 300) comparing differences between individuals in a group of A. frenatus for each acoustic feature. n, number of pulses analyzed.