Table 3.

Statistical models for multitrait animal models

	Random effects		Fixed effects				Covariates
Trait	Anim	Year × tank	Year	Year × stat	Year × sex × mat	Year × stat × sex × mat	Tsum (year)
Survival1	x	x	x
Survival2	x	x	x
Survival2sea	x	x		x
Weight1	x	x	x				x
Weight2	x	x			x
Weight3	x	x			x
Weight2sea	x	x				x
Deformation2	x	x	x
Cataract2	x	x	x
Flesh colour2sea	x	x				x
Entrail%2sea	x	x				x
Female maturity2	x	x	x
Male maturity2	x	x	x
Male maturity2sea	x	x		x

Model terms are: Anim = genetic effect of an individual with full pedigree; Year × tank = random interaction of birth year and family rearing tank; Year = fixed effect of birth year; Year × stat = fixed interaction of birth year and testing stations in fresh and sea water; Year × sex × mat = fixed interaction of birth year, sex and maturity; Year × stat × sex × mat = fixed interaction of birth year, station, sex and maturity; and Tsum (year) = covariate of cumulative temperature sum at date of recording, nested within birth year.