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. Author manuscript; available in PMC: 2014 Jan 7.
Published in final edited form as: Mol Pharm. 2012 Dec 24;10(1):337–352. doi: 10.1021/mp3004622

Table 3.

Tumor suppressor genes upregulated following decitabine treatment for 48 or 72 h in resistant breast cancer (MCF-7/ADR) cells.

No Gene Symbol Entrez Gene I.D Description Sub cellular location Function Comments
1 CD19 930 CD19 molecule Plasma Membrane Assembles with antigen receptor of B Lymphocytes Expression leads to growth inhibition and reduced tumorigenicity in human multiple myeloma29
2 CSRP2 1466 Cysteine and glycine rich protein-2 Nucleus Development and cell differentiation Acts as a repressor for NF-kappa B to suppress tumorigenesis and angiogenesis30
3 CST6 1474 Cystatin E/M Extracellular, secreted protein Physiological inhibitors of lysosomal cysteine proteases Significantly reduces cell proliferation and migration31,32
4 CRMP3 10570 Dihydropyrimidi-nase-like 4 Cytoplasm Cell signaling and cytoskeleton remodeling Direct downstream target of p53 on apoptosis. Apoptosis inducible factor in response to DNA damage33
5 DUSP4/MKP2 1846 Dual specificity phosphatase 4 Nucleus Negatively regulates Mitogen activated protein (MAP) kinase Inhibits MAP kinase signaling and thereby cell proliferation. Tumor suppressor in gliomas34
6 DUSP5 1847 Dual specificity phosphatase 5 Nucleus Negatively regulates Mitogen activated protein (MAP) kinase Direct transcriptional target of p53 to deactivate MAP - kinase pathway and thereby inhibits cell proliferation35
7 ECRG4/C2orf 40 84417 Chromosome 2 open reading frame 40 Secreted protein Probable hormone Putative tumor suppressor gene in various cancers including breast cancer36
8 EGR1 1958 Early growth response 1 Nucleus Transcriptional regulator Functions as tumor suppressor via regulation of p53 in skin carcinoma (ref). Direct regulator of multiple tumor suppressors including TGFbeta1, p53, p21 and PTEN37
9 HES6 55502 Hairy and enhancer of split 6 Nucleus cell differentiation Inhibits cell proliferation through induction of p2138
10 H19 283120 HI9, imprinted maternally expressed transcript (non-protein coding) Nucleus and Cytoplasm Acts as translational repressor Functions as a tumor suppressor in various cancers39
11 IBP6 3489 Insulin like growth factor binding protein 6 Secreted Binds to IGF-I and -II with high affinity and specificity Activates programmed cell death in non-small cell lung cancer cells both in vitro and in vivo40
12 IF127 3429 Interferon, alpha-inducible protein 27 Plasma membrane and mitochondria Promotes cell death Mediates interferon induced apoptosis characterized by release of cytochrome c, and activation of BAX and caspases - 2,3,6,8 and 9. Also sensitizes cells to other stimulus such as etopside41
13 LGALS7 3963 Lectin, galactoside-binding, soluble, 7 Cytoplasm and nucleus Involved in cell- cell and/or cell-matrix interactions Suppresses angiogenesis and tumor growth42
14 LXN 56925 Latexin Cytoplasm Endogenous carboxypeptidase inhibitor Inhibits lymphoma cell proliferation and proliferation of gastric carcinoma in vitro and in vivo43,44
15 MT1F 4494 Metallothionein 1F Cytoplasm and nucleus Binds various heavy metals Putative tumor suppressor gene in colon carcinogenesis45
16 MT1G 4495 Metallothionein 1G Cytoplasm and nucleus Binds various heavy metals Reported as tumor suppressor gene in hepatocellular carcinoma46
17 p21 1026 Cyclin-dependent kinase inhibitor 1A Cytoplasm and nucleus Binds to and inhibits cyclin-dependent kinase activity Tumor suppressor, induces cell cycle arrest in cancer cells47
18 RASD1 51655 RAS, dexamethasone-induced 1 Plasma membrane Regulates cell growth and cell differentiation Inhibits cell proliferation of G418 resistant MCF 7 breast cancer cells, lung adenocarcinoma A549 and murine fibroblasts (NIH 3T3) cells in vitro48
19 SMAD3 4088 SMAD family member 3 Cytoplasm and nucleus Signal transducer and transcriptional regulator In gastric cancer cells, SMAD3 expression restored the expression of TGF-b, p21 and p15, thereby inhibits cancer growth49
20 SOCS1 8651 suppressor of cytokine signaling 1 Intracellular, nucleus Protein kinase inhibitor, negative regulator of JAK/STAT pathway Functions as tumor suppressor gene in various cancers including breast carcinomas5052
21 THY1 7070 Thy-1 cell surface antigen Plasma membrane Cell- cell adhesion and cytoskeletal organization Suppresses the growth of human ovarian cancer and nasopharyngeal carcinoma53,54