Table 2. Species frequency of occurrence inside (in) and outside (out) cushions of T. caespitosum at four elevation sites in Nubra.
Species | Family | Clona-lity | Life | 4850 m | 5000 m | 5100 m | 5250 m | |||||||||||||
form | in | out | Fp | Fa | in | out | Fp | Fa | in | out | Fp | Fa | in | out | Fp | Fa | total | |||
Poa attenuata | Poa | yes | H | 34 | 50 | *** | *** | 24 | 39 | *** | *** | 19 | 30 | ** | 30 | 41 | ** | 267 | ||
Potentilla pamirica | Rosa | no | H | 0 | 8 | ** | * | 14 | 28 | ** | ** | 18 | 26 | *** | 31 | 29 | 154 | |||
Oxytropis tatarica | Faba | no | H | 0 | 0 | 1 | 5 | 23 | 32 | * | 20 | 32 | * | * | 113 | |||||
Astragalus strictus | Faba | no | H | 30 | 45 | ** | *** | 11 | 23 | * | ** | 0 | 1 | *** | *** | 0 | 0 | 110 | ||
Astragalus confertus | Faba | no | H | 0 | 0 | 6 | 12 | * | 15 | 25 | * | 19 | 24 | * | 101 | |||||
Draba altaica | Brassica | no | Ch | 0 | 2 | 3 | 18 | *** | *** | 10 | 16 | 22 | 30 | 101 | ||||||
Leontopodium leontopodinum | Aster | yes | H | 19 | 21 | 7 | 18 | ** | *** | 4 | 2 | 4 | 4 | 79 | ||||||
Oxytropis densa | Faba | no | H | 34 | 34 | 5 | 6 | 0 | 0 | 0 | 0 | 79 | ||||||||
Artemisia minor | Aster | no | Ch | 12 | 37 | *** | *** | 11 | 16 | 0 | 0 | 0 | 0 | 76 | ||||||
Carex pseudofoetida | Cyper | yes | H | 10 | 13 | ** | 9 | 8 | 1 | 2 | 10 | 10 | 63 | |||||||
Tanacetum tibeticum | Aster | no | Ch | 5 | 9 | 4 | 11 | * | 5 | 15 | * | * | 4 | 5 | 58 | |||||
Potentilla bifurca | Rosa | yes | H | 15 | 21 | 4 | 4 | 0 | 0 | 0 | 0 | 44 | ||||||||
Lloydia serotina | Lili | yes | G | 0 | 0 | 4 | 3 | 0 | 1 | 18 | 15 | 41 | ||||||||
Elymus schrenkianus | Poa | yes | H | 8 | 7 | 0 | 0 | 5 | 4 | 1 | 0 | 25 | ||||||||
Potentilla gelida | Rosa | no | H | 0 | 0 | 6 | 18 | *** | *** | 0 | 0 | 0 | 0 | 24 | ||||||
Potentilla multifida agg. | Rosa | no | H | 4 | 10 | 1 | 2 | 0 | 0 | 0 | 0 | 17 | ||||||||
Oxytropis platysema | Faba | no | H | 0 | 0 | 7 | 7 | 0 | 0 | 1 | 1 | 16 | ||||||||
Saussurea glacialis | Aster | no | H | 0 | 0 | 1 | 0 | 4 | 1 | 5 | 5 | 16 | ||||||||
Taraxacum sp. | Aster | no | H | 0 | 0 | 3 | 10 | * | 0 | 0 | 1 | 1 | 15 | |||||||
Trisetum spicatum | Poa | yes | H | 0 | 0 | 6 | 6 | 0 | 0 | 1 | 0 | 13 | ||||||||
Oxytropis chiliophylla | Faba | no | H | 0 | 0 | 3 | 8 | * | * | 0 | 0 | 0 | 0 | 11 | ||||||
Carex sp. | Cyper | yes | H | 5 | 4 | 0 | 0 | 0 | 0 | 0 | 0 | 9 | ||||||||
Gentianella azurea | Gentiana | no | T | 1 | 4 | 0 | 0 | 0 | 0 | 0 | 4 | 9 | ||||||||
Artemisia santolinifolia | Aster | no | Ch | 0 | 8 | *** | *** | 0 | 0 | 0 | 0 | 0 | 0 | 8 | ||||||
Elymus schugnanicus | Poa | yes | H | 3 | 4 | 0 | 0 | 0 | 0 | 0 | 0 | 7 | ||||||||
Braya humilis | Brassica | no | H | 0 | 0 | 1 | 4 | 0 | 0 | 0 | 1 | 6 | ||||||||
Thalictrum alpinum | Ranuncul | yes | H | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 3 | 6 | ||||||||
Aster flaccidus | Aster | yes | H | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 3 | ||||||||
Ephedra gerardiana | Ephedra | yes | Ch | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | ||||||||
Oxytropis pusilla | Faba | no | H | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ||||||||
Saussurea gnaphalodes | Aster | no | H | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 3 | ||||||||
Urtica hyperborea | Urtica | no | H | 0 | 2 | *** | 1 | 0 | 0 | 0 | 0 | 0 | 3 | |||||||
Potentilla saundersiana | Rosa | no | H | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | ||||||||
Waldheimia tridactylites | Aster | no | H | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 2 | ||||||||
Carex borii | Cyper | yes | H | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | ||||||||
Kobresia schoenoides | Cyper | yes | H | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | ||||||||
Lomatogonium thomsonii | Gentiana | no | T | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ||||||||
Sibbaldia tetrandra | Rosa | yes | Ch | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | ||||||||
total | 182 | 285 | 5 | 7 | 134 | 248 | 8 | 9 | 106 | 155 | 2 | 6 | 171 | 210 | 1 | 3 | ||||
average richness | 2.8 | 4.3 | 2.2 | 4.1 | 1.5 | 2.3 | 2.2 | 2.8 | ||||||||||||
total richness | 14 | 21 | 24 | 22 | 11 | 12 | 18 | 19 |
Friedman-based resampling on presence/absence (Fp) and cover-abundance (Fa) data was used to define the preferred position of a species (*** P<0.001, ** P<0.01, * P<0.05). All significant preferences refer to habitat outside cushion. Also shown is potential for clonal growth and life form (H – hemicryptophyte, Ch – chamaephyte, G – geophyte, T - terophyte).