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. 2013 Jan 18;8(1):e54539. doi: 10.1371/journal.pone.0054539

Table 3. Effects on brown treecreeper behaviour on particular substrates.

Target substrate Behaviour Parameter Estimate (±s.e.) Odds ratio ?2 d.f. P
Branch Foraging Fixed effects
• Vegetation 19.72 2 <0.001
Medium 0.61 (±0.27) 1.84
Low −0.68 (±0.28) 0.51
• Nest box 3.94 1 0.047
With boxes −0.73 (±0.37) 0.48
• CWD site 0.82 1 0.365
• Constant −1.70 (±0.27)
Random effects
• Group + σ2 = 0.14 0.080
Group/BirdID σ2 = 0.00
Branch Vigilance Fixed effects
• Vegetation 11.67 2 0.003
Medium 0.42 (±0.34) 1.52
Low −0.71 (±0.33) 0.49
• CWD site 4.32 1 0.038
In site −1.17 (±0.56) 0.31
• Nest box 2.20 1 0.138
• Constant −0.97 (±0.28)
Random effects
• Group + σ2 = 0.01 0.985
Group/BirdID σ2 = 0.00
Branch Resting and Fixed effects
Preening • Vegetation 6.20 2 0.045
Medium 0.85 (±0.51) 2.34
Low −0.30 (±0.47) 0.74
• CWD site 0.43 1 0.513
• Nest box 0.35 1 0.553
• Constant −1.22 (±0.46)
Random effects
• Group + σ2 = 0.00 0.985
Group/BirdID σ2 = 0.09
Ground Foraging Fixed effects 1.11 2 0.574
• Vegetation
• CWD site 3.75 1 0.053
• Nest box 0.11 1 0.746
• Constant −1.60 (±0.23)
Random effect
• Group + σ2 = 0.01 0.533
Group/BirdID σ2 = 0.07
Ground Vigilance Fixed effects
• Vegetation 0.13 2 0.938
• CWD site 0.02 1 0.883
• Nest box 0.10 1 0.756
• Constant −4.06 (±1.01)
Random effects
• Group + σ2 = 0.00 1.00
Group/BirdID σ2 = 0.00
Ground Resting and Fixed effects
Preening • Vegetation 0.05 2 0.975
• CWD site 0.00 1 1.000
• Nest box 0.00 1 1.000
• Constant −14.6 (±163)
Random effects
• Group + σ2 = 0.00 1.000
Group/BirdID σ2 = 0.00
Log Foraging Fixed effects
• CWD site 6.56 1 0.010
In site 0.98 (±0.38) 2.65
• Vegetation 0.42 2 0.813
• Nest box 0.17 1 0.678
• Constant −2.63 (±0.31)
Random effects
• Group + σ2 = 0.03 0.842
Group/BirdID σ2 = 0.00
Log Vigilance Fixed effects
• Vegetation 0.15 2 0.930
• CWD site 1.52 1 0.218
• Nest box 0.07 1 0.799
• Constant −0.84 (±0.29)
Random effects
• Group + σ2 = 0.00 0.434
Group/BirdID σ2 = 0.11
Log Resting and Fixed effects
Preening • Vegetation 3.53 2 0.171
• CWD site 0.65 1 0.421
• Nest box 0.50 1 0.481
• Constant −0.38 (±0.45)
Random effects
• Group + σ2 = 0.23 0.260
• Group/BirdID σ2 = 0.00
Trunk Foraging Fixed effects
• Vegetation 7.99 2 0.018
Medium −0.56 (±0.23) 0.57
Low −0.01 (±0.19) 0.99
• CWD site 11.68 1 <0.001
In site −1.07 (±0.31) 0.34
• Nest box 2.85 1 0.091
• Constant 0.40 (±0.19)
Random effects
• Group + σ2 = 0.00 0.083
Group/BirdID σ2 = 0.10
Trunk Vigilance Fixed effects
• Vegetation 8.96 2 0.011
Medium −0.43 (±0.30) 0.65
Low 0.39 (±1.48) 1.48
• CWD site 0.87 1 0.351
• Nest box 2.83 1 0.092
• Constant −0.51 (±0.27) 0.87 1 0.351
Random effects
• Group + σ2 = 0.00 0.647
Group/BirdID σ2 = 0.08
Trunk Resting and Fixed effects
Preening • Vegetation 1.64 2 0.441
• CWD site 0.04 1 0.844
• Nest box 0.29 1 0.590
• Constant −0.63 (±0.44)
Random effects
• Group + σ2 = 0.15 0.393
Group/BirdID σ2 = 0.00

Results of generalized linear mixed models (GLMMs) and generalized linear models (GLMs) using binomial distribution (logit-link structure) analysing the effects of (1) the level of ground vegetation cover (high, medium and low); (2) experimental coarse woody debris site (in or out of site); and (3) the presence or absence of nest boxes; on the probability of Brown Treecreeper individuals using each of the four target substrates: (1) branch, (2) ground, (3) log, and (4) trunk, whilst displaying the three types of behaviours (1) foraging, (2) vigilance, and (3) resting and preening. Group and individual bird nested within group were included as random effects in GLMMs (σ2 =  the variance of the random factor). The significant effects are shown in bold and include estimates (± s.e.) and odds ratios, which use high ground vegetation and outside of a 1 ha coarse woody debris site as the reference levels. Estimate for the constant is given from the full GLMMs. The total number of observations for each of the analyses by behaviour was: foraging: 663; vigilance: 374; and resting and preening: 155.