Table 3. Effects on brown treecreeper behaviour on particular substrates.
Target substrate | Behaviour | Parameter | Estimate (±s.e.) | Odds ratio | ?2 | d.f. | P |
Branch | Foraging | Fixed effects | |||||
• Vegetation | 19.72 | 2 | <0.001 | ||||
Medium | 0.61 (±0.27) | 1.84 | |||||
Low | −0.68 (±0.28) | 0.51 | |||||
• Nest box | 3.94 | 1 | 0.047 | ||||
With boxes | −0.73 (±0.37) | 0.48 | |||||
• CWD site | 0.82 | 1 | 0.365 | ||||
• Constant | −1.70 (±0.27) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.14 | 0.080 | |||||
Group/BirdID | σ2 = 0.00 | ||||||
Branch | Vigilance | Fixed effects | |||||
• Vegetation | 11.67 | 2 | 0.003 | ||||
Medium | 0.42 (±0.34) | 1.52 | |||||
Low | −0.71 (±0.33) | 0.49 | |||||
• CWD site | 4.32 | 1 | 0.038 | ||||
In site | −1.17 (±0.56) | 0.31 | |||||
• Nest box | 2.20 | 1 | 0.138 | ||||
• Constant | −0.97 (±0.28) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.01 | 0.985 | |||||
Group/BirdID | σ2 = 0.00 | ||||||
Branch | Resting and | Fixed effects | |||||
Preening | • Vegetation | 6.20 | 2 | 0.045 | |||
Medium | 0.85 (±0.51) | 2.34 | |||||
Low | −0.30 (±0.47) | 0.74 | |||||
• CWD site | 0.43 | 1 | 0.513 | ||||
• Nest box | 0.35 | 1 | 0.553 | ||||
• Constant | −1.22 (±0.46) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.00 | 0.985 | |||||
Group/BirdID | σ2 = 0.09 | ||||||
Ground | Foraging | Fixed effects | 1.11 | 2 | 0.574 | ||
• Vegetation | |||||||
• CWD site | 3.75 | 1 | 0.053 | ||||
• Nest box | 0.11 | 1 | 0.746 | ||||
• Constant | −1.60 (±0.23) | ||||||
Random effect | |||||||
• Group + | σ2 = 0.01 | 0.533 | |||||
Group/BirdID | σ2 = 0.07 | ||||||
Ground | Vigilance | Fixed effects | |||||
• Vegetation | 0.13 | 2 | 0.938 | ||||
• CWD site | 0.02 | 1 | 0.883 | ||||
• Nest box | 0.10 | 1 | 0.756 | ||||
• Constant | −4.06 (±1.01) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.00 | 1.00 | |||||
Group/BirdID | σ2 = 0.00 | ||||||
Ground | Resting and | Fixed effects | |||||
Preening | • Vegetation | 0.05 | 2 | 0.975 | |||
• CWD site | 0.00 | 1 | 1.000 | ||||
• Nest box | 0.00 | 1 | 1.000 | ||||
• Constant | −14.6 (±163) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.00 | 1.000 | |||||
Group/BirdID | σ2 = 0.00 | ||||||
Log | Foraging | Fixed effects | |||||
• CWD site | 6.56 | 1 | 0.010 | ||||
In site | 0.98 (±0.38) | 2.65 | |||||
• Vegetation | 0.42 | 2 | 0.813 | ||||
• Nest box | 0.17 | 1 | 0.678 | ||||
• Constant | −2.63 (±0.31) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.03 | 0.842 | |||||
Group/BirdID | σ2 = 0.00 | ||||||
Log | Vigilance | Fixed effects | |||||
• Vegetation | 0.15 | 2 | 0.930 | ||||
• CWD site | 1.52 | 1 | 0.218 | ||||
• Nest box | 0.07 | 1 | 0.799 | ||||
• Constant | −0.84 (±0.29) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.00 | 0.434 | |||||
Group/BirdID | σ2 = 0.11 | ||||||
Log | Resting and | Fixed effects | |||||
Preening | • Vegetation | 3.53 | 2 | 0.171 | |||
• CWD site | 0.65 | 1 | 0.421 | ||||
• Nest box | 0.50 | 1 | 0.481 | ||||
• Constant | −0.38 (±0.45) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.23 | 0.260 | |||||
• Group/BirdID | σ2 = 0.00 | ||||||
Trunk | Foraging | Fixed effects | |||||
• Vegetation | 7.99 | 2 | 0.018 | ||||
Medium | −0.56 (±0.23) | 0.57 | |||||
Low | −0.01 (±0.19) | 0.99 | |||||
• CWD site | 11.68 | 1 | <0.001 | ||||
In site | −1.07 (±0.31) | 0.34 | |||||
• Nest box | 2.85 | 1 | 0.091 | ||||
• Constant | 0.40 (±0.19) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.00 | 0.083 | |||||
Group/BirdID | σ2 = 0.10 | ||||||
Trunk | Vigilance | Fixed effects | |||||
• Vegetation | 8.96 | 2 | 0.011 | ||||
Medium | −0.43 (±0.30) | 0.65 | |||||
Low | 0.39 (±1.48) | 1.48 | |||||
• CWD site | 0.87 | 1 | 0.351 | ||||
• Nest box | 2.83 | 1 | 0.092 | ||||
• Constant | −0.51 (±0.27) | 0.87 | 1 | 0.351 | |||
Random effects | |||||||
• Group + | σ2 = 0.00 | 0.647 | |||||
Group/BirdID | σ2 = 0.08 | ||||||
Trunk | Resting and | Fixed effects | |||||
Preening | • Vegetation | 1.64 | 2 | 0.441 | |||
• CWD site | 0.04 | 1 | 0.844 | ||||
• Nest box | 0.29 | 1 | 0.590 | ||||
• Constant | −0.63 (±0.44) | ||||||
Random effects | |||||||
• Group + | σ2 = 0.15 | 0.393 | |||||
Group/BirdID | σ2 = 0.00 |
Results of generalized linear mixed models (GLMMs) and generalized linear models (GLMs) using binomial distribution (logit-link structure) analysing the effects of (1) the level of ground vegetation cover (high, medium and low); (2) experimental coarse woody debris site (in or out of site); and (3) the presence or absence of nest boxes; on the probability of Brown Treecreeper individuals using each of the four target substrates: (1) branch, (2) ground, (3) log, and (4) trunk, whilst displaying the three types of behaviours (1) foraging, (2) vigilance, and (3) resting and preening. Group and individual bird nested within group were included as random effects in GLMMs (σ2 = the variance of the random factor). The significant effects are shown in bold and include estimates (± s.e.) and odds ratios, which use high ground vegetation and outside of a 1 ha coarse woody debris site as the reference levels. Estimate for the constant is given from the full GLMMs. The total number of observations for each of the analyses by behaviour was: foraging: 663; vigilance: 374; and resting and preening: 155.