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. 2013 Jan 28;8(1):e55009. doi: 10.1371/journal.pone.0055009

Table 9. Nichols’ [41] forces estimated by the seven methods.

Method Forces
Inheritance Borrowing Substratum Selection
CM + ? + ? − ? ±
CC + ? + ? − ? ±
CR + ? + ? − ? ±
D +++ + +
M +++ ±
P1 + + − − − ? − ? +
P2 + + − − − ? − ? +
W +++ ±

A subjective view on what combination of “forces” each method estimates. Cells represent approximate effects: + (postive effect), − (negative effect), ? (unclear), ± (could be positive or negative).