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G-tetrad stacking: (A) G-tetrads have a polarity as defined by direction of the Hoogsteen hydrogen bond donor-to-acceptor pattern. Tetrad stacking can be primarily described as (B) opposite-polarity or (C) same-polarity stacking. Model stacked G-tetrad geometries used in this study varied by (D) relative rotation angle (θtet) and (E) G-tetrad separation distance (dtet).
