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The fitness effects of mutator-dependent mutations. (a) Beneficial mutations in rpoS, mglD and malT from the evolved clone (BW3767) were transferred to the ancestor (BW2952) and vice versa as shown (see §2 for details). (b) Fitness (means ± s.e.m.) of the resulting strains was measured against a reference strain containing a metC::Tn10 countable marker as described by Maharjan et al. [12].
