Table 1. Model comparison.

Model	Interaction zone		/radians					-value	bits
Null	None	−7.5	N/A	N/A	N/A	N/A	N/A	0	−69036
MF	Global	−7.5	N/A	N/A	0.76	N/A	N/A		−57976
T	K nearest-neighbours	−7.5	N/A	5	0.77	N/A	N/A	0.077	−58114
S1	Spatial, symmetric	−7.5	0.20	N/A	1.35	N/A	N/A		−60035
S2	Spatial, forward	−7.5	0.16	N/A	1.37	N/A	N/A	0	−59102
S3	Spatial, symmetric	−7.5	0.20	N/A	1.72	0.23	N/A		−62297
S4	Spatial, forward	−7.5	0.19	N/A	1.69	0.52	N/A	0	−62004
D1	Spatial, symmetric	−7.5	0.18	N/A	1.08	N/A	0.87	0.097	−58094
D2	Spatial, forward	−7.5	0.19	N/A	0.75	N/A	0.94	0.15	−58499
D3	Spatial, symmetric	−7.5	0.19	N/A	0.99	−3.59	0.92	0.17	−57963
D4	Spatial, forward	−7.5	0.19	N/A	1.08	0.32	0.92		−58512

The interaction zone structure of each model, along with the (maximum a posteri) inferred values of model parameters, the -value indicating quality-of-fit between experimental results and model simulations and the log marginal likelihood () of the model calculated from the fine scale dynamics (as shown in Figure 3. -values reported as zero are smaller than numerical precision, i.e ). N/A indicates that the model does not include the indicated parameter. The interaction zone indicates whether prawns interact with others in a spatial zone around themselves, which may be oriented either forwards or symmetrically around their centre, or with their nearest-neighbours or globally with all other individuals. Reported parameters are: , the baseline direction-change intensity; , the interaction radius for spatial models; , the number of interacting nearest-neighbours for topological models; and , the strength of interaction with negative and positive prawns respectively; and , the decay factor determining how long interaction effects persist.