Table 1.

Current classification and some properties of Grapevine leafroll-associated viruses (GLRaVs).

Virus	Genus	Coat protein (kDa)	Genome size (nt)	GenBank access. No.	ORFs	Vectors	First record fide [Boscia et al. (1995), Martelli et al. (2012)]
GLRaV-1	Ampelovirus	34	18659	JQ023131	9	Mealybugs and soft scale insects	Gugerli et al. (1984)
GLRaV-2	Closterovirus	22	16494	AY88162	8	Unknown	Zimmermann et al. (1990)
GLRaV-3	Ampelovirus	35	18498	EU259806	12	Mealybugs, soft scale and scale insects	Zee et al. (1987)
GLRaV-4	Ampelovirus	35	13830	FJ467503	6	Mealybugs	Hu et al. (1990)
GLRaV-5a	Ampelovirus	35	13384b	FR822696	6	Mealybugs	Walter and Zimmermann (1991), Zimmermann et al. (1990)
GLRaV-6a	Ampelovirus	35	13807	FJ467504	6	Mealybugs	Gugerli and Ramel (1993), Gugerli et al. (1997)
GLRaV-7	Velarivirusc	37	16496	HE588185	10	Unknown	Choueiri et al. (1996)
GLRaV-9a	Ampelovirus	35	12588b	AY29781	6	Mealybugs	Alkowni et al. (2004)
GLRaV-Pra	Ampelovirus	30	13696	AM182328	6	Mealybugs	Maliogka et al. (2009)
GLRaV-Cara	Ampelovirus	29	13626	FJ907331	6	Unknown	Abou Ghanem-Sabanadzovic et al. (2010)

^aFuture classification might list these as strains of GLRaV-4.

^bNearly complete sequence.

^cTentative classification.