Table 5. Mean rate (± SE) per minute of event-behaviour during confrontation of territorial males against intruders (♂♂ vs. ♂, N = 7, n = 28), between territorial males (♂♂ vs. ♂♂, N = 3, n = 12) and territorial males against females (♂♂ vs. ♀, N = 6, n = 24).

Behaviours	♂♂ vs ♂	♂♂vs ♂♂	♂♂vs ♀
Sounds	0.9±0.2	6±1.7	0.5±0.2
Bitings	1.7±0.2	0.1±0.02	0.6±0.15