Table 4. Parameters and values used in models on the three spatial scales.

Parameter	Meaning	Value of range	Justification
Local Scale
σu, σv	Standard deviation of dispersal east-west, north-south	21.1 sq-m day−1 , 10.6 sq-m day−1	Median value taken from [50] with a phototactic response to the rising and the setting of the sun
Field Scale
σu, σv	Standard deviation of dispersal east-west, north-south	211 sq-m day−1 , 106 sq-m day−1	Ten times local scale value for qualitative fit
u,v	Wind-shift distance east-west, north-south	104–1088 m day−1	Found by Equation 3 with parameters f, w, L fitted by genetic algorithm from the landscape scale (below)
Landscape Scale
L	Length of time mated females fly	10.2 min day−1	[53]
f	Dimensionless scaling factor	1	Fitted by genetic algorithm
wu, wv	Half-hourly mean wind speed east-westnorth-south	0.2–1.8 m s−1	Recorded at release site, but only applied when between 6.30 am and 5∶00 pm and <2.2 m s−1 as fitted by genetic algorithm