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| hFSHR | hFSH | hLHR | hLH |
| 50E | 42P | 55Y | 89R |
| 55L | 89S | 101L | 97P |
| 101R | 90D | 103Q | 98K |
| 104K | 95T | 178Y | 105D |
| 179K | 96V | 199E | - |
| 221V | 97R | 54A | - |
| 52R | 103Y | 128C | - |
| 54V | - | - | - |
| 103E | - | - | - |
The second-shell residues are italicized.
