Table 2. Summary of general linear models of the relationships between segregating load and drift load with expected heterozygosity (H _E), reflecting long-term population size and (ln-transformed) average density of A. lyrata, reflecting intraspecific competition.

Dependent variable	Segregating load (δ)						Drift load/heterosis (H)
	Intercept		HE		Density		Intercept		HE		Density
	Estimate	t	Range	F	Range	F	Estimate	t	Range	F	Range	F
Summary performance measures
Multipl. perform. I	0.19	3.21**	0.15/0.21	0.09	0.15/0.22	0.11	0.49	3.20**	2.01/−0.39	20.16**	1.39/−0.46	9.89*
ln Multipl. perform. II	0.18	3.83**	0.13/0.21	0.23	0.11/0.24	0.55	0.36	1.99(*)	1.75/−0.44	11.89**	1.21/−0.52	6.16*
Life stage components
ln Seed size	−0.01	−0.50	−0.01/−0.01	0.00	−0.01/−0.01	0.00	0.01	0.61	0.05/−0.02	2.26	0.01/0.00	0.02
Germination	0.01	0.68	0.03/0.00	0.35	−0.02/0.04	0.86	0.03	2.81*	0.05/0.02	0.50	0.08/−0.02	6.46*
Pathogen resistance	−0.03	−1.24	−0.09/0.00	1.17	−0.04/−0.02	0.07	0.06	1.60	0.26/−0.06	7.04*	0.17/−0.06	3.01
Flowering time 2009	−0.20	−1.00	−0.20/−0.20	0.00	−0.13/−0.27	0.03	0.02	0.12	0.34/−0.17	1.04	0.15/−0.12	0.23
Flowers 2009	0.14	3.41**	0.27/0.07	2.06	0.17/0.11	0.14	0.06	0.84	0.41/−0.14	5.13*	0.15/−0.04	0.49
Fruits 2009	0.15	1.94(*)	0.26/0.08	0.43	0.26/0.03	0.60	0.16	2.34*	0.40/0.02	2.53	0.11/0.21	0.17
Pollen number	0.09	2.79*	0.14/0.07	0.41	0.07/0.12	0.15	−0.06	−1.76	−0.01/−0.09	0.51	−0.03/−0.09	0.21
Pollen size	−0.01	−2.14(*)	−0.02/−0.01	0.34	−0.01/−0.01	0.08	−0.01	−1.79	−0.01/−0.01	0.05	−0.01/−0.01	0.02
Flowering time 2010	0.18	4.15**	0.47/0.01	9.58*	0.29/0.05	2.24	0.71	1.50	4.40/−1.41	12.34**	2.74/−1.42	5.24*
Flowers up to 2010	0.15	2.59*	0.19/0.13	0.07	0.20/0.10	0.17	0.39	2.48*	1.42/−0.21	9.10*	1.00/−0.26	4.51(*)
ln Fruits up to 2010	0.18	3.46**	0.18/0.19	0.00	0.19/0.17	0.01	0.23	1.95(*)	0.87/−0.14	6.03*	0.61/−0.17	2.96
Flowers up to 2012	0.16	2.46*	0.19/0.15	0.02	0.18/0.14	0.03	0.42	2.99*	1.66/−0.30	16.13**	1.11/−0.31	7.16*
ln Fruits up to 2012	0.17	3.42**	0.15/0.18	0.03	0.17/0.17	0.00	0.23	2.09(*)	0.92/−0.16	7.65*	0.62/−0.17	3.40(*)

Segregating load (δ=inbreeding depression due to one generation of selfing) and heterosis (H) were calculated as described in Materials and methods, and their relation with expected heterozygosity and A. lyrata density in nature was tested with a general linear model on population estimates (N=13). Test statistics are t-value for the intercept and F-value for the two centered covariates. The range of segregating and drift loads corresponds to model-estimated loads based on the largest and smallest observed population-level values of expected heterozygosity and A. lyrata density. Significance is indicated in bold: (*)P<0.1, *P<0.05, **P<0.01, ***P<0.001.