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Determinants for rhomboid substrate specificity. P1 and P4 are the first and fourth residues on the N-terminal side of the scissile bond, respectively, and P2′ is the second residue on the C-terminal side. The S1 and S2′ subsites of rhomboid recognize the P1 and P2′ residues of the substrate. HB, helix-breaker residue.
