Variation of reproductive success among six O. heldreichii populations throughout eastern Crete in comparison with the two study populations from Prina (black) and Orino (light grey). Total number of flowers in each population: Gournia (N = 76), Orino (N = 36), Zaros (N = 126), Kalamafka (N = 37), Meseleri (N = 61), Kroistas (N = 46), Gergeri (N = 54) and Prina (N = 80). a Male reproductive success; b Female reproductive success