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. Author manuscript; available in PMC: 2014 Jun 1.
Published in final edited form as: Ann N Y Acad Sci. 2013 May 3;1288(1):17–35. doi: 10.1111/nyas.12109

Table 4.

Task-specific handedness in captive chimpanzees

Measure #R #A #L HI s.e. t-value
Throwing (n = 89) 50 16 23 .30 .091 3.81**
Gesture (n = 347) 169 133 45 .26 .025 10.17**
TUBE (n= 542) 272 118 152 .14 .024 5.69**
Ball (n = 54) 31 14 9 .28 .083 3.59 **
Reaching (n = 345) 105 161 79 .03 .022 1.49
Tool Use (n =192) 63 52 74 −.06 .040 −1.42
Coconut (n = 157) 61 32 64 −.01 .054 −0.23
Bimanual Feeding (n = 185) 73 65 47 .08 .032 2.34 *
(T) Grooming (n = 200) 48 49 103 .03 .032 0.95
(B) Grooming (n= 125) 28 84 13 .13 .042 2.95 **
(U) Grooming (n = 178) 52 78 48 -.01 .036 −0.17
*

p < .05

**

p < .01.

(T) reflects total frequency in right and left hand use for grooming, (B) bimanual grooming, (U) unimanual grooming. #R = number or right-handed, #A = number of ambiguously-handed, #L = number of left-handed. Hand preference classification were based on z-scores calculated on the basis of frequencies in left and right hand use for each task. Subjects with z-scores ≥ 1.96 were classified as right-handed, those with z-scores ≤ −1.96 were classified as left-handed. All others were classified as ambiguously-handed.