Table 2. Phenotypic characterization of Nissle 1917 and its isogenic Csr mutants.

Strain	Glycogen	Motility	Biofilm
wild-type	++	++	+
ΔcsrA51	++++	+	+++
ΔcsrB	+	++	+
ΔcsrC	+	+++	+
ΔcsrBC	+	+++	++
ΔcsrD	++	+	+

Each experiment was performed in three independent biological replicates. The data shown here are semi-quantitative.