FIGURE 2.
(a) Examples of alternative male morphs in Onthophagus taurus (top) and O. nigriventris (bottom). Large males are shown on the left and small males on the right. Note that females (not shown) are entirely hornless in both species. (b) Rare reversed sexual dimorphism in O. sagittarius. Males (shown on left) also lost ancestral male dimorphism. (Reprinted with permission from Ref 17. Copyright 2012 Hindawi Publishers)
