Within- and between-cross variation in (A) SR length, (B) time to female sperm ejection after the end of copulation, (C) the number of first-male sperm still in storage at the time of remating, and (D) the proportion of second-male sperm among all sperm stored (i.e., S2). Each point represents an individual isoline cross (for simplicity, the reciprocal crosses are combined by nuclear genotype); error bars indicate SE. For statistics on heritability, see Table 1.