The biosynthetic pathway of carotenoids in plants. GGPP, Geranylgeranyl diphosphate; PSY, phytoene synthase; PDS, phytoene desaturase; ZDS, ζ-carotene desaturase; CRTISO, carotene isomerase; LCYB, lycopene β-cyclase; CYP97A3 and CYP97C1, cytochrome P450 enzymes; NSY, neoxanthin synthase; LCYE, lycopene ε-cyclase; CRTR-E, ε-carotene hydroxylase. Enzymes in red were examined in this study.