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B5 SCR4 is critical for actin-tail formation by superinfecting EEV. EEV preparations from VACV mutants lacking B5 SCR1–4 (vSCR0), 2–4 (vSCR1) or 4 (vSCR1–3) or with a P189S mutation in B5 SCR4 (vB5P189S) were added to RK13 A33–A36-4 cells for 30 min. Cells were then fixed and stained with phalloidin and an anti-F13 mAb to visualize actin tails and EEV particles. The results are expressed in number of actin tails detected per 106 virus particles and represent the mean±sd, n = 3.
