Table 1.

Parameters measured for organism, segments, and muscles

Variable description	Variable name
Whole-organism behavioral
Head position	r(Head)
Centroid position	r(Centroid)
Tail position	r(Tail)
Segmental movement
Mouth hook position	r(Mouth Hooks)
Tail position	r(A8)
Gut position	r(Gut)
Segment position	r(T2, T3, A1-A7)
Individual muscle contractions
Length of muscle DO1 on segments A2–A7	l(DO1)
Length of muscle LT1 on segments A2–A7	l(LT1)
Length of muscle VL1 on segments A2–A7	l(VL1)