Table 1. Partial fitnesses and pairwise interaction rates with cliquing behavior.

Phenotype	Partial fitnesses		Interaction rates		Total fitnesses
	Bards	Poets	Bards	Poets
Bards	1	1 − cs	f + (1 − f)q2	(1 − f)(p2 + 2pq)	1 − cs(1 − f)(p2 + 2pq)
Poets	1 − cs	1 + s	(1− f)q2	f + (1 − f)(p + 2pq)	1 + s − (1 + c)s(1 − f)q2

Bards are recessive for ancestral allele BARD. Parameter s ≥ 0 is the selective advantage of two poets communicating with one another, and 0 ≤ c ≤ 1/s is the relative penalty of cross communication. The strength of cliquing behavior is represented by 0 ≤ f ≤ 1. The alleles POET and BARD have frequencies p and q = 1 − p in the population, respectively