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. 2013 Jul 23;8(7):e70038. doi: 10.1371/journal.pone.0070038

Table 2. Abiotic predictor variables grouped in four categories (COVER, RESISTANCE, SOIL and STRUCTURE) and measured at different spatial scales.

Category Scale Abbreviation Description Unit
Percentage cover (COVER) 25×25 cm quadrat, 50×50 cm quadrat, 1×1 m quadrat BARESOIL_25, BARESOIL_50, BARESOIL_1 Bare soil cover %
CALLUNA_25, CALLUNA_50, CALLUNA_1 Calluna vulgaris cover %
DUNG_25, DUNG_50, DUNG_1 Cattle dung cover %
GRAMINOID_25, GRAMINOID_50, GRAMINOID_1 Graminoid species cover %
HERB_25, HERB_50, HERB_1 Total herb species (including graminoids) cover %
LICHEN_25, LICHEN_50, LICHEN_1 Lichen cover %
LITTER_25, LITTER_50, LITTER_1 Leaf litter cover %
MOSS_25, MOSS_50, MOSS_1 Moss cover %
ROOT_25, ROOT_50, ROOT_1 Root (alive and dead) cover %
SEDUM_25, SEDUM_50, SEDUM_1 Sedum album cover %
SHRUB_25, SHRUB_50, SHRUB_1 Total shrub species (including Calluna vulgaris, Vaccinium myrtillus and V. uliginosum) cover %
STONE_25, STONE_50, STONE_1 Stone cover %
VACCINIUM_25, VACCINIUM_50, VACCINIUM_1 Vaccinium myrtillus and V. uliginosum cover %
WOOD_25, WOOD_50, WOOD_1 Dead wood cover %
Vegetation resistance (RESISTANCE) 1×1 m quadrat COVER_N, COVER_E, COVER_S, COVER_W Vegetation cover estimated in a 6×6 square grid (0.13 m2) vertically centered in the quadrat at each cardinal direction %
HEIGHT Mean (N = 7) Calluna vulgaris height cm
SQUARE_N, SQUARE_E, SQUARE_ S, SQUARE_W Number of squares crossed by vegetation in a 6×6 square grid (0.13 m2) vertically centered in the quadrat at each cardinal direction
Soil features (SOIL) 1×1 m quadrat LITTER_DEPTH Mean (N = 5) depth of the leaf litter layer cm
SOIL_0.063 Soil (horizon A) particle size <0.063 mm %
SOIL_0.125 Soil (horizon A) particle size 0.063–0.125 mm %
SOIL_0.25 Soil (horizon A) particle size 0.125–0.25 mm %
SOIL_0.50 Soil (horizon A) particle size 0.25–0.50 mm %
SOIL_1 Soil (horizon A) particle size 0.50–1 mm %
SOIL_2 Soil (horizon A) particle size 1–2 mm %
SOIL_HUMID Soil (horizon A) humidity %
SOIL_OM Soil (horizon A) organic matter content %
SOIL_PH Soil (horizon A) pH
Habitat structure (STRUCTURE) 3 m radius, 6 m radius OPEN_3, OPEN_6 Number of bare ground patches
SHRUB_3, SHRUB_6 Number of shrubs >1 m height
STRUCTURE_3, STRUCTURE_6 Habitat structure classes: 1) 100% closed vegetation (Calluna vulgaris), 2) few distant open bare ground patches, 3) 50% open - 50% closed patches, 4) mostly open patches, 5) 100% open patches, 6) 100% closed vegetation (shrub species >1 m height)
6 m radius DIST_OPEN Distance to the nearest bare ground patch cm
DIST_SHRUB Distance to the nearest shrub >1 m height cm