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Disrupted morphogenetic patterning of trabecular and compact myocardium. In situ hybridization analysis of Hey2 and Tbx20 expression; both Hey2 and Tbx20 expression are significantly higher in compact myocardium of wild-type hearts, but are expanded to trabecular myocardium in FKBP12 mutant hearts. Red arrows denote trabecular myocardium; blue arrows denote compact wall.
