Table 1. List of abbreviations used in this paper.

Abbreviation	Description
C	Control treatment
CDX	Treatment to detect epigenetic effects; males with a Control X-chromosome that have been produced by DX females
DX	Females with a double X-chromosome (i.e. two X-chromosomes attached at the centromere)
GO	Gene Ontology database
LHm	The outbred laboratory stock used in these experiments
MLX	Male-limited X-chromosome evolution treatment (and males derived from this treatment)
RB	Recombination box; accessory population to MLX populations allowing recombination between X-chromosomes
SA	Sexually antagonistic, here used in the context of sexually antagonistic zygotic drive