(A) In a temporal- or spatial-specific way, protein poly(ADP-ribosyl)ation is reversed by PARG activity. Therefore, HRP proteins are not poly(ADP-ribosyl)ated and will bind to their target pre-mRNAs for RNA processing or translational control, which further regulates developmental patterns. (B) Once PARG activity is downregulated in different tissues or developmental stages, HRP proteins are poly(ADP-ribosyl)ated, which inhibits Hrps from binding to pre-mRNA. Therefore, the pre-mRNA bound by Hrps are not processed or translated, and developmental patterns are changed. (C) Mutating HRP proteins blocks pre-mRNA processing or translation.